This paper identifies multiple mechanisms: as well as psbA gene mutation there is a non-target site resistance mechanism of enhanced metabolism. Of course, we expect multiple mechanisms of herbicide resistance as evident here in both target site psbA gene mutation and non-target site enhanced metabolism resistance.
Wild radish (Raphanus raphanistrum) is a globally important weed of crops. Two atrazine-resistant wild radish populations (R1 and R2), collected from the Western Australia grain belt, were investigated for resistance to photosystem II (PSII) herbicides.