The obvious evolutionary reality is that persistent use of harvest weed seed control (HWSC) is a selection pressure for any mechanisms enabling L. rigidum seed to avoid HWSC. For example, seed shatters before grain harvest or a greater percentage of retained seed at a height below that at which the crop is cut in the harvesting operation.
Seed production of annual weeds persisting through cropping phases replenishes/establishes viable seed banks from which these weeds will continue to interfere with crop production. Harvest weed seed control (HWSC) systems are now viewed as an effective means of interrupting this process by targeting mature weed seed, preventing seed bank inputs.